Saban said he would play OE. I am taking this to mean "outside end". Basically this is a OLB that walks up to the line, outside of the Defensive End. I think you will see Jason Taylor play the other OE. With Kevin Carter and Vonnie Holliday at DE's and Larry Chester at NT. Seau and Thomas will start the season at ILB and they will slowly bring in Crowder for Seau.
The best article to see a diagram of this is one at Phins.com called Nick Saban's 3-4 Defense. Its a great article.